General rule of thumb for plant spacing is to take into consideration the full size of the plant being grown. That is how big it gets when it's full grown, then space that distance apart. Closer spacing could require excessive pruning to compensate for crowding, and/or lead to inadequate air movement between plants and foliage. Inadequate air movement can also lead to fungal diseased, as well as pest infestations.
Another problem with crowding inside a greenhouse with inadequate air movement is rapid c02 depletion. The plants will still use the same amount of c02 weather they are crowded or not, but if there is inadequate air movement between the plants and foliage, the c02 the plants use cant be replenished fast enough through air circulation. Then the plants growth will slow or even stop, they simply cant grow without c02. Plants get all their carbon (c02) from the atmosphere, and carbon is the basic building block for all plant and animal life.
P.S.
You can help air circulation by using fans, especially oscillating fans that move back and forth.
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